THE FORMATION OF VEGETABLE MOULD THROUGH THE ACTION OF WORMS
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wissenschaft. Zoologie,’ B. xix., 1869, p. 611.
{18a} For instance, Mr. Bridgman and Mr. Newman (‘The Zoologist,’ vol.
vii. 1849, p. 2576), and some friends who observed worms for me.
{18b} ‘Familie der Regenwürmer,’ 1845, p. 18.
{31} ‘The Zoologist,’ vol. vii. 1849, p. 2576.
{32} ‘Familie der Regenwürmer,’ p. 13. Dr. Sturtevant states in the
‘New York Weekly Tribune’ (May 19, 1880) that he kept three worms in a
pot, which was allowed to become extremely dry; and these worms were
found “all entwined together, forming a round mass and in good
condition.”
{33} ‘De Lumbrici terrestris Hist. Nat.’ p. 19.
{34} ‘Archives de Zoologie expérimentale,’ tom. vii. 1878, p. 394. When
I wrote the above passage, I was not aware that Krukenberg
(‘Untersuchungen a. d. physiol. Inst. d. Univ. Heidelberg,’ Bd. ii. p.
37, 1877) had previously investigated the digestive juice of Lumbricus.
He states that it contains a peptic, and diastatic, as well as a tryptic
ferment.
{35a} On the action of the pancreatic ferment, see ‘A Text-Book of
Physiology,’ by Michael Foster, 2nd edit. pp. 198–203. 1878.
{35b} Schmulewitsch, ‘Action des Sucs digestifs sur la Cellulose.’
Bull. de l’Acad. Imp. de St. Pétersbourg, tom. xxv. p. 549. 1879.
{40} Claparède doubts whether saliva is secreted by worms: see
‘Zeitschrift für wissenschaft. Zoologie,’ B. xix. 1869, p. 601.
{41a} Perrier, ‘Archives de Zoolog. expér.’ July, 1874, pp. 416, 419.
{41b} ‘Zeitschrift für wissenschaft. Zoologie,’ B. xix, 1869, pp.
603–606.
{46} De Vries, ‘Landwirth. Jahrbücher,’ 1881, p. 77.
{49} M. Foster, ‘A Text-Book of Physiology,’ 2nd edit. 1878, p. 243.
{50} M. Foster, _ut sup._ p. 200.
{53} Claparède remarks (‘Zeitschrift für wisseuschaft. Zoolog.’ B. 19,
1869, p. 602) that the pharynx appears from its structure to be adapted
for suction.
{58} An account of her observations is given in the ‘Gardeners’
Chronicle,’ March 28th, 1868, p. 324.
{59a} London’s ‘Gard. Mag.’ xvii. p. 216, as quoted in the ‘Catalogue of
the British Museum Worms,’ 1865, p. 327.
{59b} ‘Familie der Regenwürmer,’ p. 19.
{79} In these narrow triangles the apical angle is 9° 34′, and the basal
angles 85° 13′. In the broader triangles the apical angle is 19° 10′ and
the basal angles 80° 25′.
{89a} See his interesting work, ‘Souvenirs entomologiques,’ 1879, pp.
168–177.
{89b} Möbius, ‘Die Bewegungen der Thiere,’ &c., 1873, p. 111.
{90} ‘Annals and Mag. of N. History,’ series ii. vol. ix. 1852, p. 333.
{93} ‘Archives de Zoolog. expér.’ tom. iii. 1874, p. 405.
{97} I state this on the authority of Semper, ‘Reisen im Archipel der
Philippinen,’ Th. ii. 1877, p. 30.
{101} Dr. King gave me some worms collected near Nice, which, as he
believes, had constructed these castings. They were sent to M. Perrier,
who with great kindness examined and named them for me: they consisted of
_Perichæta affinis_, a native of Cochin China and of the Philippines; _P.
Luzonica_, a native of Luzon in the Philippines; and _P. Houlleti_, which
lives near Calcutta. M. Perrier informs me that species of Perichæta
have been naturalized in the gardens near Montpellier and in Algiers.
Before I had any reason to suspect that the tower-like castings from Nice
had been formed by worms not endemic in the country, I was greatly
surprised to see how closely they resembled castings sent to me from near
Calcutta, where it is known that species of Perichæta abound.
{102} ‘Zeitschrift für wissenschaft. Zoolog.’ B. xxviii. 1877, p. 364.
{108} ‘Zeitschrift für wissenschaft. Zoolog.’ B. xxviii. 1877, p. 356.
{113} Perrier, ‘Archives de Zoolog. expér.’ tom. 3, p. 378, 1874.
{126} This case is given in a postscript to my paper in the ‘Transact.
Geolog. Soc.’ (Vol. v. p. 505), and contains a serious error, as in the
account received I mistook the figure 30 for 80. The tenant, moreover,
formerly said that he had marled the field thirty years before, but was
now positive that this was done in 1809, that is twenty-eight years
before the first examination of the field by my friend. The error, as
far as the figure 80 is concerned, was corrected in an article by me, in
the ‘Gardeners’ Chronicle,’ 1844, p. 218.
{128} These pits or pipes are still in process of formation. During the
last forty years I have seen or heard of five cases, in which a circular
space, several feet in diameter, suddenly fell in, leaving on the field
an open hole with perpendicular sides, some feet in depth. This occurred
in one of my own fields, whilst it was being rolled, and the hinder
quarters of the shaft horse fell in; two or three cart-loads of rubbish
were required to fill up the hole. The subsidence occurred where there
was a broad depression, as if the surface had fallen in at several former
periods. I heard of a hole which must have been suddenly formed at the
bottom of a small shallow pool, where sheep had been washed during many
years, and into which a man thus occupied fell to his great terror. The
rain-water over this whole district sinks perpendicularly into the
ground, but the chalk is more porous in certain places than in others.
Thus the drainage from the overlying clay is directed to certain points,
where a greater amount of calcareous matter is dissolved than elsewhere.
Even narrow open channels are sometimes formed in the solid chalk. As
the chalk is slowly dissolved over the whole country, but more in some
parts than in others, the undissolved residue—that is the overlying mass
of red clay with flints,—likewise sinks slowly down, and tends to fill up
the pipes or cavities. But the upper part of the red clay holds
together, aided probably by the roots of plants, for a longer time than
the lower parts, and thus forms a roof, which sooner or later falls in,
as in the above mentioned five cases. The downward movement of the clay
may be compared with that of a glacier, but is incomparably slower; and
this movement accounts for a singular fact, namely, that the much
elongated flints which are embedded in the chalk in a nearly horizontal
position, are commonly found standing nearly or quite upright in the red
clay. This fact is so common that the workmen assured me that this was
their natural position. I roughly measured one which stood vertically,
and it was of the same length and of the same relative thickness as one
of my arms. These elongated flints must get placed in their upright
position, on the same principle that a trunk of a tree left on a glacier
assumes a position parallel to the line of motion. The flints in the
clay which form almost half its bulk, are very often broken, though not
rolled or abraded; and this may be accounted for by their mutual
pressure, whilst the whole mass is subsiding. I may add that the chalk
here appears to have been originally covered in parts by a thin bed of
fine sand with some perfectly rounded flint pebbles, probably of Tertiary
age; for such sand often partly fills up the deeper pits or cavities in
the chalk.
{131} S. W. Johnson, ‘How Crops Feed,’ 1870, p. 139.
{136a} ‘Nature,’ November 1877, p. 28.
{136b} ‘Proc. Phil. Soc.’ of Manchester, 1877, p. 247.
{138a} ‘Trans. of the New Zealand Institute,’ vol. xii., 1880, p. 152.
{138b} Mr. Lindsay Carnagie, in a letter (June 1838) to Sir C. Lyell,
remarks that Scotch farmers are afraid of putting lime on ploughed land
until just before it is laid down for pasture, from a belief that it has
some tendency to sink. He adds: “Some years since, in autumn, I laid
lime on an oat-stubble and ploughed it down; thus bringing it into
immediate contact with the dead vegetable matter, and securing its
thorough mixture through the means of all the subsequent operations of
fallow. In consequence of the above prejudice, I was considered to have
committed a great fault; but the result was eminently successful, and the
practice was _partially_ followed. By means of Mr. Darwin’s
observations, I think the prejudice will be removed.”
{139} This conclusion, which, as we shall immediately see, is fully
justified, is of some little importance, as the so-called bench-stones,
which surveyors fix in the ground as a record of their levels, may in
time become false standards. My son Horace intends at some future period
to ascertain how far this has occurred.
{147} Mr. R. Mallet remarks (‘Quarterly Journal of Geolog. Soc.’ vol.
xxxiii., 1877, p. 745) that “the extent to which the ground beneath the
foundations of ponderous architectural structures, such as cathedral
towers, has been known to become compressed, is as remarkable as it is
instructive and curious. The amount of depression in some cases may be
measured by feet.” He instances the Tower of Pisa, but adds that it was
founded on “dense clay.”
{148} ‘Zeitschrift für wissensch. Zoolog.’ Bd. xxviii., 1877, p. 360.
{149} See Mr. Dancer’s paper in ‘Proc. Phil. Soc. of Manchester,’ 1877,
p. 248.
{166a} ‘Leçons de Géologie pratique,’ 1845, p. 142.
{166b} A short account of this discovery was published in ‘The Times’ of
January 2, 1878; and a fuller account in ‘The Builder,’ January 5, 1878.
{183} Several accounts of these ruins have been published; the best is
by Mr. James Farrer in ‘Proc. Soc. of Antiquaries of Scotland,’ vol. vi.,
Part II., 1867, p. 278. Also J. W. Grover, ‘Journal of the British Arch.
Assoc.’ June 1866. Professor Buckman has likewise published a pamphlet,
‘Notes on the Roman Villa at Chedworth,’ 2nd edit. 1873 Cirencester.
{187} These details are taken from the ‘Penny Cyclopædia,’ article
Hampshire.
{210} “On the denudation of South Wales,” &c., ‘Memoirs of the
Geological Survey of Great Britain,’ vol. 1., p. 297, 1846.
{211} ‘Geological Magazine,’ October and November, 1867, vol. iv. pp.
447 and 483. Copious references on the subject are given in this
remarkable memoir.
{212} A. Tylor “On changes of the sea-level,” &c., ‘ Philosophical Mag.’
(Ser. 4th) vol. v., 1853, p. 258. Archibald Geikie, Transactions Geolog.
Soc. of Glasgow, vol. iii., p. 153 (read March, 1868). Croll “On
Geological Time,” ‘Philosophical Mag.,’ May, August, and November, 1868.
See also Croll, ‘Climate and Time,’ 1875, Chap. XX. For some recent
information on the amount of sediment brought down by rivers, see
‘Nature,’ Sept. 23rd, 1880. Mr. T. Mellard Reade has published some
interesting articles on the astonishing amount of matter brought down in
solution by rivers. See Address, Geolog. Soc., Liverpool, 1876–77.
{213} “An account of the fine dust which often falls on Vessels in the
Atlantic Ocean,” Proc. Geolog. Soc. of London, June 4th, 1845.
{215} For La Plata, see my ‘Journal of Researches,’ during the voyage of
the _Beagle_, 1845, p. 133. Élie de Beaumont has given (‘Leçons de
Géolog. pratique,’ tom. I. 1845, p. 183) an excellent account of the
enormous quantity of dust which is transported in some countries. I
cannot but think that Mr. Proctor has somewhat exaggerated (‘Pleasant
Ways in Science,’ 1879, p. 379) the agency of dust in a humid country
like Great Britain. James Geikie has given (‘Prehistoric Europe,’ 1880,
p. 165) a full abstract of Richthofen’s views, which, however, he
disputes.
{217a} These statements are taken from Hensen in ‘Zeitschrift für
wissenschaft. Zoologie.’ Bd. xxviii., 1877, p. 360. Those with respect
to peat are taken from Mr. A. A. Julien in ‘Proc. American Assoc.
Science,’ 1879, p. 354.
{217b} I have given some facts on the climate necessary or favourable
for the formation of peat, in my ‘Journal of Researches,’ 1845, p. 287.
{220} A. A. Julien “On the Geological action of the Humus-acids,” ‘Proc.
American Assoc. Science,’ vol. xxviii., 1879, p. 311. Also on “Chemical
erosion on Mountain Summits;” ‘New York Academy of Sciences,’ Oct. 14,
1878, as quoted in the ‘American Naturalist.’ See also, on this subject,
S. W. Johnson, ‘How Crops Feed,’ 1870, p. 138.
{222} See, for references on this subject, S. W. Johnson, ‘How Crops
Feed,’ 1870, p. 326.
{223} This statement is taken from Mr. Julien, ‘Proc. American Assoc.
Science,’ vol. xxviii., 1879, p. 330.
{224a} The preservative power of a layer of mould and turf is often
shown by the perfect state of the glacial scratches on rocks when first
uncovered. Mr. J. Geikie maintains, in his last very interesting work
(‘Prehistoric Europe,’ 1881), that the more perfect scratches are
probably due to the last access of cold and increase of ice, during the
long-continued, intermittent glacial period.
{224b} Many geologists have felt much surprise at the complete
disappearance of flints over wide and nearly level areas, from which the
chalk has been removed by subaerial denudation. But the surface of every
flint is coated by an opaque modified layer, which will just yield to a
steel point, whilst the freshly fractured, translucent surface will not
thus yield. The removal by atmospheric agencies of the outer modified
surfaces of freely exposed flints, though no doubt excessively slow,
together with the modification travelling inwards, will, as may be
suspected, ultimately lead to their complete disintegration,
notwithstanding that they appear to be so extremely durable.
{225a} ‘Archives de Zoolog. expér.’ tom. iii. 1874, p. 409.
{225b} ‘Nouvelles Archives du Muséum,’ tom. viii. 1872, pp. 95, 131.
{226} Morren, in speaking of the earth in the alimentary canals of
worms, says, “præsepè cum lapillis commixtam vidi:” ‘De Lumbrici
terrestris Hist. Nat.’ &c., 1829, p. 16.
{227} Perrier, ‘Archives de Zoolog. expér.’ tom. iii. 1874, p. 419.
{228a} Morren, ‘De Lumbrici terrestris Hist. Nat.’ &c., p. 16.
{228b} ‘Archives de Zoolog. expér.’ tom. iii. 1874, p. 418.
{234} This conclusion reminds me of the vast amount of extremely fine
chalky mud which is found within the lagoons of many atolls, where the
sea is tranquil and waves cannot triturate the blocks of coral. This mud
must, as I believe (‘The Structure and Distribution of Coral-Reefs,’ 2nd
edit. 1874, p. 19), be attributed to the innumerable annelids and other
animals which burrow into the dead coral, and to the fishes,
Holothurians, &c., which browse on the living corals.
{236} Anniversary Address: ‘The Quarterly Journal of the Geological
Soc.’ May 1880, p. 59.
{244} Mr. James Wallace has pointed out that it is necessary to take
into consideration the possibility of burrows being made at right angles
to the surface instead of vertically down, in which case the lateral
displacement of the soil would be increased.
{259} ‘Elements of Geology,’ 1865, p. 20.
{265} ‘Leçons de Géologie pratique, 1845; cinquième Leçon. All Élie de
Beaumont’s arguments are admirably controverted by Prof. A. Geikie in his
essay in Transact. Geolog. Soc. of Glasgow, vol. iii. p. 153, 1868.
{266} ‘Illustrations of the Huttonian Theory of the Earth,’ p. 107.
{269} Mr. E. Tylor in his Presidential address (‘Journal of the
Anthropological Institute,’ May 1880, p. 451) remarks: “It appears from
several papers of the Berlin Society as to the German ‘high-fields’ or
‘heathen-fields’ (Hochäcker, and Heidenäcker) that they correspond much
in their situation on hills and wastes with the ‘elf-furrows’ of
Scotland, which popular mythology accounts for by the story of the fields
having been put under a Papal interdict, so that people took to
cultivating the hills. There seems reason to suppose that, like the
tilled plots in the Swedish forest which tradition ascribes to the old
‘hackers,’ the German heathen-fields represent tillage by an ancient and
barbaric population.”
{284} White of Selborne has some good remarks on the service performed
by worms in loosening, &c., the soil. Edit, by L. Jenyns, 1843, p. 281.
{285} ‘Zeitschrift für wissenschaft. Zoolog.’ B. xxviii. 1877, p. 360.
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