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Nitrogen
Nitrogen is very important to insure the growth of vine without
which a good crop may not be expected. Lands vary more widely in
nitrogen content than in phosphorus and potash. Sandy soils are
commonly deficient in this element and often difficult to keep
supplied. Here liberal applications are needed. Up to a hundred
pounds[5] of actual nitrogen may prove profitable where other
conditions justify. Heavier soils, well managed and manured during
rotation, require less nitrogen and fair results may be obtained
with no fertilizer where investment must be kept to a minimum.
Form of nitrogen to be used is largely a matter of economy though
nitrate for part of it may be desirable early in the season when
soil is cold and nitrification slow. Nitrate is desirable for side
dressing but even here ammonia and other forms are now considered
suitable when the soil is warm.
Failure to Set Fruit
Why do tomatoes sometimes run to vine with failure to set fruit?
This is an old, old query and, since 1918, has been, directly or
indirectly, the occasion of more research projects than any other
horticultural topic. Kraus and Kraybill[6] set the ball a-rolling
with a paper which called attention first to the observations of
Klebs in Germany in which he emphasized the fact that external
conditions influence conditions within the plant which in turn
influence performance--a veritable chain of causation. Kraus and
Kraybill then undertook to relate performance (vegetative growth and
fruitfulness) to internal conditions, chiefly carbohydrate and
nitrogen content of the plant tissues. These, in turn, were traced
back to treatments applied to the soil.
They suggested four combinations of vegetation and fruitfulness in
plants as follows:
1. Non-vegetative and non-fruitful. Plants whose carbohydrate
supply has been cut off, say by removal of leaves which make
carbohydrates. These plants were low in carbohydrate and high
in nitrogen.
2. Vegetative and non-fruitful. These plants were well
supplied with both carbohydrates and nitrogen. They were of
the sort we describe as having "run to vine."
3. Vegetative and fruitful. These plants were well supplied
with carbohydrates, but not so liberally supplied with
nitrogen, thus, providing a balance between the two that was
favorable for a good crop.
4. Non-vegetative and non-fruitful. These plants had ample
opportunity for carbohydrate making, but were underfed with
nitrogen and so could not perform well in either vegetation or
fruit-making.
Kraus and Kraybill conclude that there are certain balances between
these two groups of compounds--nitrogenous and carbohydrate--which
determine the nature of the plant's performance--whether there will
be too little vegetative growth to permit a crop, whether the plants
will "run to vine" or whether they will show good growth of both
foliage and fruit.
From experiments in the same field, using definite amounts of
nitrate of soda per plant, Work[7] concluded that while adequate
carbohydrate supply is necessary for fruiting, excess carbohydrate
did not, in itself, occasion unfruitfulness but was more likely to
represent an accumulation of material unused by reason of deficiency
in some other factor--often nitrogen.
It was shown that nitrate of soda does not injure tomatoes until a
concentration in the soil is attained which is strong enough to
plasmolyze the cells, that is to withdraw water from them by
osmosis. Nor were a wide variety of nitrogen and moisture and manure
treatments sufficient to induce the Bonny Best variety to "run to
vine." Some varieties are subject to this trouble, mostly of the
large, late types.
Murneek[8] has shown that the fruitfulness of a plant may greatly
affect its internal condition, its vegetative performance and its
later setting of fruit. A heavy load of developing fruit, with
limited soil resources, tends to limit growth and setting. Removal
of fruit induces renewal of vegetative growth and of fruit setting.
Failure to set fruit favors vigorous vine growth. This failure may
be traceable to various causes. (1) To damage to floral parts as the
blasting of the pistil by heat and drouth. Flowers of some varieties
show tendency toward elongation of pistils with subsequent failure
to develop normal fruit. Smith and Howlett have shown that
environmental conditions as well as heredity influence this
elongation. (2) To injury by insects as thrips. (3) To the character
of the variety used, the Bonny group being very slightly susceptible
to failure from over feeding with nitrogen while some late sorts
readily "run to vine." (4) Shortage of nutrient elements as nitrogen
or phosphorus or others. (5) Lack of adequate light or short day. In
such cases, there may be excess of nitrogen for current need with
resultant over-development of leafage. Thus, excess vegetative
growth may be a result as well as a cause of poor setting.
Phosphorus
Fertilizer experiments fairly generally point to the frequency with
which phosphorus is the limiting factor among nutrients in tomato
production. MacGillivray[9] has studied the phosphorus content of the
various parts of the plant, concluding that this element is important
throughout and not alone in seed making or in rapidly growing parts as
has been believed. Hepler and Kraybill[10] found some years ago and
others more recently have confirmed the influence of liberal phosphorus
treatments upon earliness.
[Illustration: FIGURE 5.--Effect of omission of phosphorus from complete
fertilizer in Western New York.]
Potash
The potash requirement of the tomato has not been as thoroughly
studied as the requirement for the other two major elements. It is
thought that potash has a part in building up sugars into more
complex carbohydrates.
The consensus of fertilizer experiments suggests that potash is less
important on most soils than phosphorus and nitrogen but that if
these elements are in good supply, increased yields from potash are
likely.
Lanham in Texas was unable to find a relation between potash
fertilization and resistance to shipping hazards.
Stable Manure
Stable manure has long been recognized as useful for tomatoes. It is
generally considered better to apply it to the preceding crop or at
least the preceding fall than to use it just before setting of
plants. If spring application is necessary, it is better to use well
rotted manure. Stable manure is low in phosphorus. An approximate
statement would be that 10 tons of manure is roughly equivalent to
one ton of a 6-3-6 fertilizer. Thus, 1,000 pounds of 18%
superphosphate would bring the analysis to 6-12-6 which would be
generally regarded as a good balance.
A recent publication[11] from Pennsylvania emphasizes the value of
manures and of phosphorus.
Placement and Side Dressing
Recent experiments have shown the desirability of placing fertilizer
close to but not in contact with the roots of the young plants. When
newly set and before new roots have developed is the time when
nutrient material close at hand is needed to give the plant a
vigorous send-off. Transplanters have been devised with attachments
to place the fertilizer in bands at each side of the row of tomatoes
and about two inches deep.
Recent experiments, notably by Sayre[12] of New York, have shown the
advantage of dissolving fertilizer materials in the water used for
transplanting tomatoes. One combination of materials consists of
ammo-phos, 14-48, 2 parts and potassium nitrate, 1 part. Five to
eight pounds of this mixture are dissolved in 50 gallons of water
and about 1/4 pint or 1/2 cup is applied to each plant, usually by
the transplanting machine. There are other suitable mixtures of
nutrients for this purpose. A very small investment in starter
solutions has shown material increase in total yield. The practice
places immediately available nutrients in the soil at the time and
place to be of maximum usefulness to plants that have been severely
root-pruned and have not yet had opportunity to rebuild the root
system.
[Illustration:
_Courtesy Campbell Soup Co._
FIGURE 6.--Cultivating and side-dressing tomatoes.]
Another critical stage in tomato growth comes when much fruit has
been set in the clusters and demands upon plant and soil are
especially heavy. At this stage, side dressing with nitrogen is
helpful in maintaining plant growth and providing resources for
growth and maturing of fruit. On sandy or nutrient-deficient soils,
more than one side dressing may be advisable. Sodium nitrate is
commonly used but other materials are suitable after the soil has
warmed up. Side dressing with fertilizer in solution has been
recommended recently by Tiedjens of New Jersey.
III
THE BEST IN SEED IS NONE TOO GOOD
A tomato crop may be much poorer than the seed from which it grows
but it can be no better.
The tomato seed is short-oval and flattened in shape, covered
thickly with short silky hairs. The embryo or baby plant is coiled
in a spiral and imbedded in the endosperm (reserve food supply).
Three or four years is generally given as the life of the seed but
it often remains viable much longer--up to 10 or 12 years in extreme
cases. Good seed should germinate 85% to 90%.
Tomato seed sprouts readily, requiring fairly warm temperature, say,
70 deg. to 75 deg. F. for best results. It germinates very slowly at 40 deg. to
50 deg. F.
Breeding
Being a major vegetable crop, the tomato has received much attention
from plant breeders. Objectives sought include good cannery type,
resistance to the fusarium wilt and other diseases, better
greenhouse forms, improved general market and home garden sorts, and
varieties adapted for arduous conditions such as hot and dry summers
or very short growing seasons.
The tomato is largely but not wholly self pollinated and pollen is
not carried far. Thus, it is not difficult to breed to practically a
pure-line condition.
Tomatoes for seed are usually ground up and the seed and fine pulp
are separated from the skins and coarse material by screening. The
juice, fine pulp and seeds are allowed to ferment from 24 to 48
hours, or until the jelly-like pulp is readily washed away. After
washing, the seed is dried in thin layers and stored. A bushel of
tomatoes may be expected to yield 2-1/2 to 4 ounces of seed and an
acre of tomatoes, from 100 to 225 pounds. These vary greatly
according to varieties and conditions.
Wellington[13] and others have shown that first generation seed from
crosses of suitable varieties show a marked increase of vigor
(heterosis or hybrid vigor) over either parent or over the later
generations. This fact would seem to offer possibilities in
practical use, but it has not thus far proved of value.
Selection Methods
Many growers find it profitable to save their own tomato seed. The
plant is an annual, the important characters are quite readily
observed and natural crossing is not serious. For these reasons, the
enterprise is not as difficult as with most vegetables, although, if
done well, it makes heavy demands in labor and care at a time when
the grower has much else to do.
The first step in selection is to establish clearly the ideal to be
sought, recording it in detail on paper for future reference.
Selections should be made on the basis of the plant, not of the
individual fruit. It is the plant that is reproduced and the seed
from "crown clusters" is no earlier than seed from later settings.
The field should be searched soon after blooming time and plants
that appear promising should be marked. These plants should be
examined three or four times as the season advances, and markers
pulled from plants that do not measure up to the desired standard.
Suppose ten plants remain; all fruits from each of these may be
saved, keeping the seed of each plant separate. All or part of the
seed may be planted in separate rows the next year for further
selection and to note which parents best transmit their excellent
points. If only a small amount of seed is required, direct
selections may be made for use in planting for the general crop. If
a larger amount of seed is required, seed from one or two of the
best plants should be planted in multiplication plats. Off-type
plants should be removed from such plantings, but otherwise all the
seed may be saved for use. Repeated selection results in constant
improvement until the stock becomes a "pure line" or practically so.
Lindstrom of Iowa has led in research on the genetics of tomatoes,
chromosome relations and mode of inheritance. Many scientific papers
deal with inheritance methods and results. The Yearbook of
Agriculture (U.S.D.A.) for 1937 contains a valuable chapter on
tomato breeding. It may also be had as Yearbook Separate 1581.
Certification
As with certain other kinds of seeds, certification service for tomatoes
has now been set up in several states. Certification is a most useful
incentive toward care in breeding and handling and affords valuable
assurance to the buyer. It is necessary to know just what is guaranteed
by the certificate. It is at the same time wise to be informed as to the
inclusiveness and methods of the certification.
The Ideal Variety
In breeding for better varieties of tomatoes, the following are some of
the characters to be sought:
(1) A vigorous vine which is necessary to produce abundant
fruit and to protect from sunscald.
(2) Resistance to disease especially to fusarium.
(3) High productiveness with moderate number of fruits per
cluster--say, 5 to 8.
[Illustration: FIGURE 7.--Types of tomato interiors. 1, 5. Small fruited
sorts. 2, 6, 9. Bonny Best. 3, 7, 10. Chalk Jewel. 3, 7, 11. Stone. 4,
8. Earliana. 12. Ponderosa.]
(4) Evenness of maturity. This is somewhat out of line with
the nature of the tomato but much could be accomplished toward
the goal of varieties that make their crop and are gone,
eliminating long picking periods and the drag of inferior
fruit toward the end of the season. The so-called
determinate habit of some varieties such as Pritchard is a
step in this direction.
(5) Size suitable for expected use and for market demand.
Greenhouse tomatoes are generally smaller than those for
cannery. Uniformity of size is increasingly important with
wide-spread use of the lug-box pack and of small consumer
cartons.
(6) Globular to oblong shape is desirable for market but is
less important for cannery. Form should be symmetrical, even
and smooth.
(7) Color should be deep and rich, fully and evenly developed,
inside and out. Red is generally preferred to pink. The
difference between red and pink tomatoes does not reside in
the flesh but in the presence of yellow pigment in the skin of
the former while the skin of the latter is without pigment.
Yellow tomatoes are also extant.
[Illustration: FIGURE 8.--The Earliana tomato. A picture of a single
fruit cannot adequately describe a variety. 1-3. Rough types, common in
older strains. 4. Typical interior. 5, 6. Stem end. 7-9. Good type
resulting from selection. 10-12. Pointed-round type occurring frequently
in improved strains. A. Unusually large cluster. B. Typical Earliana
cluster showing compound branching. C. Unbranched cluster of Bonny Best
for comparison.]
(8) Skin should be thick and tough. This safeguards against
damage on the way to market and favors ease of peeling. Those
saladists who serve sliced tomatoes with skins unremoved, may
call for a thin, tender skin but this practice finds no
encouragement from discriminating partakers.
(9) Flesh should be abundant in thick walls with a minimum of
watery pulp surrounding seeds. In general, a structure of many
small cells is desirable.
Varieties
_Earliana._--The earliness of this old and popular variety outweighs
its demerits where this character is required. The past ten years
have seen material improvement.
Earliana is early, of small vine, with small leaves and leaflets.
Clusters are compoundly branched, with many fruits. The fruits are
of medium size, deep oblate, cross section often elliptical rather
than circular. There are many rough irregular fruits, varying in
this respect with breeding and conditions of growth. Color is red,
not too deep and tending to be poorly developed at the stem end.
Interior consists of many small cells with thin walls.
_Bison._--represents a group of varieties bred for rigorous climates
of our most northerly states. A. F. Yeager formerly of North Dakota,
later of Michigan, now of New Hampshire has led in this
development.
_Victor._--is a new variety bred originally by Yeager but introduced
by K. C. Barrons of Michigan. It affords smoother, deeper and better
colored fruits about as early as Earliana. It is determinate in
habit and shy in foliage, increasing danger of sunscald. Rich soil
and ample moisture are needed for its best development. _Bounty_ and
_Home Garden_ are similar.
_Penn State._--Penn State, developed by C. E. Myers of Pennsylvania,
is not as early as Earliana. It is similar in fruit characters
though distinctly better in color and shape. It is marked by short
branches (determinate habit) and is designed to give an early crop
to be followed by prompt abandonment of the planting. It is not to
be confused with Penn State Earliana.
_Bonny Group._--This group embraces our leading second early
varieties widely used for home garden, greenhouse, market and
cannery in the north. It includes _Bonny Best_, _John Baer_ and
_Chalk Jewel_ with many additional names and with much confusion of
characters among them.
Bonny Best is second early and of medium plant growth. Fruits are
deep oblate to flattened globe, even and smooth, of good red color,
with few large, thick-walled cells.
[Illustration: FIGURE 9.--Marglobe plant.]
Varieties and strains of this group vary in growth and yield, in
size, shape and earliness of fruit and in suitability for
greenhouse, market, cannery and juice. _Stokesdale_ and _Scarlet
Dawn_ are meritorious newer names in the group.
_Marglobe._--This variety was developed by the late Dr. F. J.
Pritchard from a cross between Marvel, a French variety lending
resistance to fusarium and Globe, an old variety of fine size and
shape. It is widely used, north and south, for market--green or
ripe, for cannery and to some extent, for forcing.
[Illustration: FIGURE 10.--Marglobe fruit.]
Marglobe is a midseason variety, with large vine and foliage,
resistant to fusarium and nailhead spot. Fruits are nearly globular,
shapely and smooth, medium to large, scarlet red, with medium number
of thick walled cells. Marglobe is rather subject to deep radial
cracks.
_Pritchard_ is of the general type of Marglobe but is earlier, with
short branching habit and resistance to nailhead rust and to
fusarium, and, perhaps, is less subject to cracking.
_Greater Baltimore_ is used chiefly for canning in long-season
districts. It is late, with large vine, large flat fruits of
excellent scarlet red, outside and in, with many thick walled cells.
_Indiana Baltimore_ is a variant widely grown in the mid-west for
cannery.
_Rutgers_ was developed by L. G. Schermerhorn at the New Jersey
Experiment Station for fine juice and canning characters--color,
flavor and substance. Growth is vigorous and yields are heavy;
fruits are large, flattened and well colored.
_Gulf State Market_ is a second early shipping tomato, generally
harvested green. It is flattened in shape, of well developed pink
color and good interior.
_Comet Group._--These trace mostly to English or other European
origin and are increasingly used for greenhouse and for staking
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